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CO2 activation and conversion by the CODH/ACS enzymatic complex


EMSL Project ID
51914

Abstract

The purpose of the proposal is to computationally characterize the key biochemical and biophysical features of carbon monoxide dehydrogenase (CODH) and acetyl-coenzyme A synthase (ACS) enzymatic complex. This complex catalyzes the conversion of CO2 to CO and the subsequent condensation of CO with a methyl intermediate and coenzyme A to form acetyl-coenzyme A. We aim to understand the factors regulating substrate binding, CO2 and CO activation, and reaction pathways. These include metal coordination chemistry, active site residues directly interacting with the catalytic center, and ultimately the entire protein.

The theoretical investigation of the CO2 and CO activation by the CODH/ACS complex requires the ability to obtain reliable structural, spectroscopic, electrochemical, reactivity and free energy information for systems of high complexity. We will obtain this information using long-timescale molecular dynamics simulations, and large-scale QM/MM free energy simulations. Specific issues we will address: (i) the driving force for CO/CO2 binding to the Ni centers; (ii) the correlation between the metal centers oxidation states and CODH/ACS chemistry; (iii) the role of the interactions at the active site during C-O and C-C bond breaking or formation; (iv) the structural and electronic determinants that modulate the redox properties of the catalytic metal clusters; (v) the role of the metal clusters flexibility and protein dynamics. Hypotheses elaborated on the basis of our calculations will be tested against experimental assays performed by Ragsdale at the University of Michigan Ann Arbor.

Project Details

Project type
Large-Scale EMSL Research
Start Date
2021-10-01
End Date
2023-10-01
Status
Closed

Team

Principal Investigator

Simone Raugei
Institution
Pacific Northwest National Laboratory

Co-Investigator(s)

Bojana Ginovska
Institution
Pacific Northwest National Laboratory

Team Members

Maureen Kitheka
Institution
Pacific Northwest National Laboratory

Busra Dereli Caglayan
Institution
Pacific Northwest National Laboratory

Suman Samantray
Institution
Pacific Northwest National Laboratory

Jack Fuller
Institution
Pacific Northwest National Laboratory

Conrad Johnston
Institution
Pacific Northwest National Laboratory

Hoshin Kim
Institution
Pacific Northwest National Laboratory

Qi Huang
Institution
Pacific Northwest National Laboratory

Stephen Ragsdale
Institution
University of Michigan